the albino lemma gene is a pleiotropic gene in dependent of the black lemma gene, the black lemma gene has epistatic dominance over the albino lemma gene.

• 白化颖壳基因具有一因多效性；该基因与黑颖壳基因为相互独立非等位基因，黑颖壳基因对该基因有显性上位作用。