They are Rhtls on chromosome 4B, Rht8 on 2D, Rht9 on 7B, and Rht (Wan), a new GA-insensitive dwarfing gene on chromosome 4D produced from mutation proba-bly.

• Rht8和Rht9分别位于染色体2D和7B上; 一对可能通过诱变产生的对赤霉酸不敏感的矮秆基因；暂命名为Rht(Wan)；位于染色体4D上.