The wall formation conforms with the basic type of Davis. It consists of an epidermis, an endothecium, 3-4 middle layers of cells and 2-3 layers of secretory tapetum.

• 多个孢原起源于花药原基表皮之下,花药壁的发育属于Davis的基本型,药壁由7或9层细胞组成,即表皮、药室内壁、3或4层中间层、2或3层分泌型绒毡层。