The total storage of C, N, P, K, Ca, Mg, Mn, Al and Fe in the CWD was 48699 t/hm 2, but return of elements to the forest ground in the litterfall only made up 7 77% of the nutrient storage of the CWD.

• 粗死木质物是森林中重要的养分库，其中的C、N、P、K、Ca、Mg、Mn、Al和Fe元素的总贮量达到48699 t/hm 2，每年通过凋落物方式积累于林地表的养分元素只占其总量的7.；77%25