The main results are as follows: (1) Anther wall comprises epidermis, fibrous endothecium, one middle layer and glandular tapetum with two-nucleate cells.

• 主要结果如下：（1）花药壁由四层细胞构成，由外到内分别为表皮、药室内壁、中层和绒毡层,花药壁发育方式为双子叶型。