The epidermis at ventral side of dorsal microphylls and dorsal side of ventral microphylls are similar, which are sub-square or rectangular in shape and straight or sinuolate in anticlinal walls.

• 背叶腹面与腹叶背面的表皮细胞形态相似，近方形或矩形，垂周壁纹饰平直或浅波浪形。