The development of research has resolved in part many important problems, such as the origin and molecular mechanism of ovule ontogenesis.But these problems still require further study.

• 近年来，随着研究手段的不断改进，研究者对许多关键性的问题，如胚珠的起源、胚珠发生发育的分子机制已经有了一定的认识，但是这些问题并未得到圆满的解决。