The degenerating tapetum nuclei in the middle of anther locules are from the tapetum cells, which undergo mitosis, then intrude into the anther locules and degenerate in situ at the early stage.

• 绒毡层异型起源，属于腺质型绒毡层，药室内具有的退化绒毡层核是早期该层细胞有丝分裂凸入药室中央并原位退化形成的；