Stems erect, climbing, arching, or prostrate, glabrous or hairy, usually with prickles or bristles, sometimes with glandular hairs, rarely unarmed.

• 茎直立，攀援，拱起，或平卧，无毛或有毛，通常具皮刺或刚毛，有时具腺毛，很少无刺。