Males mark by spraying a combination of urine and scent from two glands at the base of their tales. While marking they scrape the ground,often leaving a noticeable depression which itself may serve as a signal to other lions.

• 雄狮主要通过散布尿和尾巴后部两个腺体发出来的混合气味，来确定自己的领地范围，通常这也是对其他狮子的一个有效警示。