It improves during the second stage in plant history with the origin in the Siluoian Period of seedless vascular land plants. These plants proliferated in the coal forests of the late Palaeozoic Era.

• 志留纪开始了植物演化史的第二个阶段，此时出现了无种籽的陆地维管植物，在晚古生代，这些植物在生成煤的森林中得到迅速繁殖。