At maturation stage, the plastoglobules released from elaioplasts coalesced with unsaturated lipids from endoplasmic reticulum to form the pollenkitt mass deposited on the exine.

• 花粉粒双细胞晚期，营养层细胞成熟，造油体中之脂肪球与内质网产生之脂肪颗粒瓦解共同形成花粉鞘附著于花粉外壁上。