And although they never increased the lake salinity to levels of the open ocean, they would still have given oceangoing animals a means for penetrating the heart of Amazonia.

• 虽然内陆湖泊的盐度并没有因此升高到开阔海域的程度，但这些海水应该还是提供了海栖动物深入亚马逊河流域中心的一些途径。